| 1 | Aberdeen Group plc | 7,966,498 | $13.0M | 0.02% | Share |
| 2 | Opaleye Management Inc. | 6,789,738 | $11.1M | 1.46% | Share |
| 3 | NORTHWESTERN MUTUAL WEALTH MANAGEMENT CO | 33,982 | $9.5M | 0.01% | Share |
| 4 | VANGUARD GROUP INC | 4,356,647 | $7.1M | 0.00% | Share |
| 5 | Nantahala Capital Management, LLC | 3,990,169 | $6.5M | 0.21% | Share |
| 6 | Telemark Asset Management, LLC | 3,846,411 | $6.3M | 0.63% | Share |
| 7 | BlackRock, Inc. | 3,734,525 | $6.0M | 0.00% | Share |
| 8 | AMERIPRISE FINANCIAL INC | 3,793,574 | $6.0M | 0.00% | Share |
| 9 | MILLENNIUM MANAGEMENT LLC | 3,135,192 | $5.0M | 0.00% | Share |
| 10 | BlackRock, Inc. | 2,374,556 | $3.0M | 0.00% | Share |
| 11 | STATE STREET CORP | 1,603,510 | $2.0M | 0.00% | Share |
| 12 | GEODE CAPITAL MANAGEMENT, LLC | 1,755,731 | $2.0M | 0.00% | Share |
| 13 | UBS Group AG | 805,785 | $1.3M | 0.00% | Share |
| 14 | BlackRock, Inc. | 663,869 | $1.0M | 0.00% | Share |
| 15 | Private Advisor Group, LLC | 654,581 | $1.0M | 0.00% | Share |
| 16 | FRANKLIN RESOURCES INC | 740,010 | $1.0M | 0.00% | Share |
| 17 | NORTHERN TRUST CORP | 631,714 | $1.0M | 0.00% | Share |
| 18 | JACOBS LEVY EQUITY MANAGEMENT, INC | 1,135,190 | $1.0M | 0.00% | Share |
| 19 | RENAISSANCE TECHNOLOGIES LLC | 737,485 | $1.0M | 0.00% | Share |
| 20 | Assenagon Asset Management S.A. | 566,547 | $923K | 0.00% | Share |
| 21 | VANGUARD GROUP INC | 552,293 | $900K | 0.00% | Share |
| 22 | PERKINS CAPITAL MANAGEMENT INC | 529,500 | $863K | 0.69% | Share |
| 23 | Post Resch Tallon Group Inc. | 511,748 | $834K | 0.70% | Share |
| 24 | Potomac Capital Management, Inc. | 300,000 | $489K | 0.39% | Share |
| 25 | Stonepine Capital Management, LLC | 256,106 | $417K | 0.30% | Share |
| 26 | Invesco Ltd. | 149,983 | $244K | 0.00% | Share |
| 27 | Diametric Capital, LP | 112,254 | $182K | 0.05% | Share |
| 28 | STRS OHIO | 99,100 | $161K | 0.00% | Share |
| 29 | SG Americas Securities, LLC | 50,290 | $82K | 0.00% | Share |
| 30 | PARADIGM CAPITAL MANAGEMENT INC/NY | 50,000 | $81K | 0.00% | Share |
| 31 | GROUP ONE TRADING LLC | 35,700 | $58K | 0.00% | Call |
| 32 | VANGUARD GROUP INC | 32,602 | $53K | 0.00% | Share |
| 33 | EVERGREEN CAPITAL MANAGEMENT LLC | 29,000 | $47K | 0.00% | Share |
| 34 | Krilogy Financial LLC | 30,101 | $46K | 0.00% | Share |
| 35 | Persistent Asset Partners Ltd | 27,813 | $45K | 0.02% | Share |
| 36 | Elmwood Wealth Management, Inc. | 21,695 | $35K | 0.01% | Share |
| 37 | AIGEN INVESTMENT MANAGEMENT, LP | 17,629 | $28K | 0.01% | Share |
| 38 | State of Wyoming | 16,870 | $27K | 0.00% | Share |
| 39 | China Universal Asset Management Co., Ltd. | 14,718 | $23K | 0.00% | Share |
| 40 | Invesco Ltd. | 13,011 | $21K | 0.00% | Share |
| 41 | Invesco Ltd. | 11,596 | $18K | 0.00% | Share |
| 42 | GAMCO INVESTORS, INC. ET AL | 10,000 | $16K | 0.00% | Share |
| 43 | Caitong International Asset Management Co., Ltd | 9,395 | $15K | 0.04% | Share |
| 44 | Mirae Asset Global Investments Co., Ltd. | 4,389 | $7K | 0.00% | Share |
| 45 | UBS Group AG | 3,734 | $6K | 0.00% | Share |
| 46 | Torren Management, LLC | 3,801 | $6K | 0.00% | Share |
| 47 | PRESCOTT GROUP CAPITAL MANAGEMENT, L.L.C. | 2,568,359 | $4K | 0.37% | Share |
| 48 | GAMMA Investing LLC | 2,604 | $4K | 0.00% | Share |
| 49 | CWM, LLC | 1,794 | $3K | 0.00% | Share |
| 50 | Zurcher Kantonalbank (Zurich Cantonalbank) | 1,587 | $2K | 0.00% | Share |